This reference list is not intended to be exhaustive but gives the references quoted in the text on other pages, plus many other key references to the literature. References are grouped under the following headings:

Distribution and conservation of D. plantarius

Dispersal and locomotion in Dolomedes species

Diet and prey-capture in Dolomedes species

Population biology of Dolomedes species

Taxonomy, systematics and genetics of the genus Dolomedes

Distribution and conservation of other Dolomedes species 

Literature on other Pisaurids

Key general references for spider identification and general biology


  Distribution and conservation of D. plantarius

Clark, M., & Clark, D., 2003  Baseline survey of the fen raft spider Dolomedes plantarius in Glamorgan. CCW Contract Science Number 603. Countryside Council for Wales, Glamorgan.

Clark, M. J. 2004,  Field notes 2003/04 for Tennant Canal and Crymlyn Bog – Dolomedes
plantarius personal observations. Unpublished to the Countryside Council for Wales, Glamorgan.


Duffey E. 1958   Dolomedes plantarius Clerk, a spider new to Britain, found in the upper Waveney Valley, Transactions of the Norfolk and Norwich Naturalist's Society 18 (7), pp.1-5.

Duffey E. 1995  The distribution, status and habitats of Dolomedes fimbriatus (Clerk) and D.plantarius (Clerk) in Europe. In: Ružicka V. (Ed.), Proceedings of the 15th European Colloquium of Arachnology, Institute of Entomology, Ceské Budejovice, pp.54-65.

Gajdoš P., Toft S., Langemark S., and Scharff N. 2000  The fishing spiders Dolomedes (Araneae, Pisauridae) in Denmark, Steenstrupia 25 (2), pp.201-208.

Gallon, R. C. 2001 Survey of the Gwent Levels wetland reserve for the fen raft spider Dolomedes plantarius. CCW Contract Science Number 489. Countryside Council for Wales, Glamorgan.

Gallon, R. 2007  Monitoring invertebrate features on SSSI's: Fen Raft Spider on Crymlyn Bog & Pant-y-Sais, West Glamorgan. CCW Contract Science Number 787. Countryside Council for Wales, Glamorgan.

Harms, D., Dunlop, J., & Schütt, K., 2009 Neue Nachweise der Gerandeten Wasserspinne Dolomedes plantarius in Brandenburg (Araneae: Pisauridae). Arachnologische Mitteilungen [online]. 37, 1-8. 

Van Helsdingen, P. 2006 Characteristic spider species of peat bog fenlands in the Netherlands (Araneae). In C. Deltschev & P. Stoev (eds.), European Arachnology 2005. Acta Zoologica Bulgarica, Supplement 1: 115–124.

Van Helsdingen, P. J. 2006  Characteristic spider species of peat bog fenlands in the Netherlands (Araneae). Acta zoologica bulgarica 2006, 115-124.

Van Helsdingen P.J. 1995 Ecology and distribution of Dolomedes in Europe (Araneida, Solomedidae), Proceedings of the 9th International Colloquium of the European Invertebrate Survey, Helsinki, 3-4 September 1993. WWF Finland 7, pp.41-46.

Holec, M., 2000 Spiders (Araneae) of the fishpond eulittoral zone. Ekológia (Bratislava) 19 (Supplement 4): 51–54.

Iorio, E. & Villepoux, O. 2012 Découverte de l’araignée Dolomedes plantarius (Clerck, 1757) (Araneae, Pisauridae) des trois nouvelles stations en Provence-Alpes-Côtes d’Azur, et réflexion sur la nécessité d’une protection régionale du genre Dolomedes. Bulletins de la Societé Linnéenne de Bordeaux 147: 43–54.

Ivanov, V., Prishepchik, O., Setrakova, E., 2017 Dolomedes plantarius (Araneae, Pisauridae) in Belarus: records, distribution and implications for conservation. Arachnology Letters [online] 54, 33-37.

Jones, E., 1992. The status of Dolomedes plantarius (Clerck) on Pevensey Levels, East Sussex in August 1991. English Nature, Peterborough.

Kirby P. 1990   Dolomedes plantarius in East Sussex, Newsletter of the British Arachnological Society 58, pp.18

Lecigne, S. 2006 Redécouverte de Dolomedes plantarius (Clerck, 1758) (Araneae, Pisauridae) en région Nord - Pas-de-Calais (France), actualisation de sa distribution en France et aperçu de la situation en Europe. Revue arachnologique, série 2, (3), 28-41.

Leroy, B., Paschetta, M., Canard, A., Bakkenes, M., Isaia, M. & Ysnel, F., 2013. First assessment of effects of global change on threatened spiders: Potential impacts on Dolomedes plantarius (Clerck) and its conservation plans. Biological Conservation [online]. 161, 155-163. 

Neet, C. & Delarze, R. 1992 Note sur la présence de Dolomedes plantarius Clerck (Araneae, Pisauridae) dans la région de Grangettes avec quelques commentaires sur son habitat. Bulletin Romand d’Entomologie 10: 81–83.

Roelofs-Ditters, E. 2003  Een nieuwe vindplaats van Dolomedes plantarius (Clerck, 1757) in Noord-Brabant. (Araneae, Pisauridae). Nieuwbrief Spined 18: 18.

RŮŽIČKA, V., 1998 New records of spiders from littorals in the Czech Republic. Arachnologische Mitteilungen 16: 1–7.

Smith H. 2000  The status and conservation of the fen raft spider (Dolomedes plantarius) at Redgrave and Lopham Fen National Nature Reserve, England, Biological Conservation 95, pp.153-164

Smith H. 2000  The status and conservation of the fen raft spider (Dolomedes plantarius) at Redgrave and Lopham Fen National Nature Reserve in 2006, Click here to Download: (764 KB).

SZINETÁR C., 2001 Somogy megye pókfaunája (Aranea). Natura Somogyiensis 1, 57-70.

Terhivuo, J. 1993 Novelties to the Finnish spider fauna (Araneae) and notes on species having bitten man. Memoranda Societatis pro Fauna et Flora Fennica 69: 53–56.

Thornhill W.A., 1985  The distribution of the great raft spider, Dolomedes plantarius, on Redgrave and Lopham fens. Transactions of the Suffolk Naturalists' Society 21: 18-22.

Trotignon, J. & Williams, T. 1994 Guide nature de la Brenne. Paris, Argenton-sur-Creusel’Association pour la Conservation du Patrimoine Natural de la Gabrière (Indre).

Unruh, M., 2008 Neue Nachweise der Listspinne Dolomedes plantarius (Clerck, 1757) im Gebiet der Mittelelbe, Sachsen-Anhalt (Araneida: Pisauridae) Hercynia N. F. 41, 143 –154

World Conservation Monitoring Centre 1996 Dolomedes plantarius. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.4. <>. Downloaded on 22 February 2011


  Dispersal and locomotion in D. plantarius

Gorb S.N. and Barth F.G. 1994  Locomotor behavior during prey-capture of a fishing spider, Dolomedes plantarius (Araneae:Araneidae): galloping and stopping, The Journal of Arachnology 22, pp.89-93

Kreiter N., & Wise D.H., 1996  Age-related changes in movement patterns in the fishing spider, Dolomedes triton (Araneae, Pisauridae). Journal of Arachnology 24: 24-33.

STRATTON, G.E., SUTER, R.B. & MILLER, P.R. 2004  Evolution of water surface locomotion by spiders: a comparative approach. Biological Journal of the Linnean Society 81, (1), 63–78.

Suter R.B. 1999  Cheap transport for fishing spiders (Araneae, Pisauridae): the physics of sailing on the water surface, Journal of Arachnology 27, pp.489-496

Suter R.B. & Gruenwald, J. 2000  Predator avoidance on the water surface? Kinematics and efficacy of vertical jumping by Dolomedes (Araneae, Pisauridae). Journal of Arachnology 28, (2), 201-210



 Diet and prey-capture in Dolomedes species

 Bleckmann H. and Lotz T. 1987 - The vertebrate-catching behaviour of the fishing spider Dolomedes triton (Araneae, Pisauridae), Animal Behaviour 35, pp.641-651

Bleckmann H. and Rovner J.S. 1984  Sensory ecology of a semi-aquatic spider (Dolomedes triton): I.Roles of vegetation and wind-generated waves in site selection, Behavioural Ecology and Sociobiology 14, 297-301.

Bleckmann H. and Barth 1984  Sensory ecology of a semi-aquatic spider (Dolomedes triton): II.The release of predatory behavior by water surface waves, Behavioural Ecology and Sociobiology 14, 303-312.

Bonnet, P., 1930  La mue, l'autotomie et la régeneration chez les araignées avec une étude des Dolomedes d'Europe. Bulletin de la Société d'histoire naturelle de Toulouse 65, 237-700.

Perevozkin1,V. P., Lukyantsev, S. V. and Gordeev, M. I. 2004  Comparative Analysis of Foraging Behavior in Aquatic and Semiaquatic Spiders of the Genera Argyroneta, Dolomedes, Pirata, and Pardosa. Journal Russian Journal of Ecology 35, (2), 103-109.

Roland C. and Rovner J.S. 1983 Chemical and vibratory communication in the aquatic pisaurid spider Dolomedes triton, Journal of Arachnology 11, 77-85.

Williams, D. S. 1979 The feeding behaviour of New Zealand Dolomedes species (Araneae:. Pisauridae). N. Z. J. Zool. 6, 95-105.

Zimmermann M., & Spence J.R., 1989  Prey use of the fishing spider Dolomedes triton (Pisauridae, Araneae): An important predator of the neuston community. Oecologia 80: 187-194.


  Population biology

Arnqvist G. 1992  Courtship behaviour and sexual cannibalism in the semi-aquatic fishing spider, Dolomedes fimbriatus (Clerk) (Araneae: Pisauridae). Journal of Arachnology 20, 222-226

Bleckmann, H. and Bender, M. 1987  Water Surface Waves Generated by the Male Pisaurid Spider Dolomedes triton (Walckenaer) during Courtship Behavior. Journal of Arachnology 15, (3), 363-369.

Johnson, J.C. 2001 Sexual cannibalism in fishing spiders (Dolomedes triton): an evaluation of two explanations for female aggression towards potential mates. Anim. Behav. 61, 905–914.

Krafft, B., & Horel, A. 1980  Comportement maternel et les relations mères-jeunes chez les araignées Reprod. Nutr. Develop 20, 747-758.

Palmgren, P., 1939  Ökologische und physiologische Untersuchungen über die Spinne Dolomedes fimbriatus (CL.). Acta Zool Fenn 24: 1-42.

Spence, J.R., Zimmermann, M., & Wojcicki, J.P., 1996  Effects of food limitation and sexual cannibalism on reproductive output of the nursery web spider Dolomedes triton (Araneae: Pisauridae). Oikos 75 (3), 373-382.

Vugdelic, M., Goodacre, S., Smith, H. & Hewitt, G., 2004 Preliminary analysis of the genetic structure in the fen spider Dolomedes plantarius (Araneae, Pisauridae). In D. V. Logunov & D. Penney (eds.), European Arachnology 2003. Arthropoda Selecta Special Issue 1: 343–348.

Zimmerman M., & Spence J.R., 1998  Phenology and life-cycle regulation of the fishing spider Dolomedes triton Walckenaer (Araneae, Pisauridae) in central Alberta. Canadian Journal of Zoology 76 (2), 295-309.

Zimmermann M., & Spence J.R., 1992  Adult population dynamics and reproductive effort of the fishing spider Dolomedes triton (Araneae, Pisauridae) in central Alberta. Canadian Journal of Zoology, 70 (11),2224-2233.



 Taxonomy and Systematics of Dolomedes

Carico, J.E., 1973. The Nearctic Species of the Genus Dolomedes (Araneae: Pisauridae). Bulletin of the Museum of Comparative Zoology 144, 435-488.

Clerck, C., 1757  Svenska spindlar, uti sina hufvud-slågter indelte samt under några och sextio särskildte arter beskrefne och med illuminerade figurer uplyste. Stockholmiae, 154 pp.

Gajdoš P., Toft S., Langemark S., and Scharff N. 2000 The fishing spiders Dolomedes (Araneae, Pisauridae) in Denmark, Steenstrupia 25 (2), 201-208.

Hahn, C. W., 1835  Die Arachniden. Nürnberg, Zweite Band, pp. 57-75.

Huber, K.C., Haider, T.S., Muller, M. W., Huber, B.A., Schweyen, R.J. & Barth, F.G. (1993)DNA Sequence Data Indicates the Polyphyly of the Family Ctenidae (Araneae). Journal of Arachnology, 21 (3), 194-201.

Ji, YJ, Smith H, Zhang, DZ & G.M. Hewitt 2004  Ten polymorphic microsatellite loci for paternity and population genetic analysis in the fen raft spider (Dolomedes plantarius) . Molecular Ecology Notes 4, 274-276.

Latreille, P. A. 1804  Tableau methodique des Insectes. Nouv. Dict. d'Hist. Nat., Paris 24, 129-295.

Müller, O. F. 1776  Zoologicae danicae prodromus, seu animalium daniae et norvegiae indigenarum, characteres, nomina et synonyma imprimis popularium. Hafniae, 282 pp. (Araneae, pp. 192-194).

Olivier G. A. 1789 Araignée, Aranea. Encycl. méth. Hist. nat. Ins. Paris 4: 173-240.

Renner, F. 1987  Revision der europäischen Dolomedes-Arten (Araneida: Pisauridae). Stuttg. Beitr. Naturk. (A) 406, 1-15.

Santos, A. J. 2007  A phylogenetic analysis of the nursery-web spider family Pisauridae, with emphasis on the genera Architis and Staberius (Araneae: Lycosoidea). Zoologica Scripta 36 (5), 489–507.

Simon, E. 1937  Les arachnides de France. Tome VI. Synopsis générale et catalogue des espèces françaises de l'ordre des Araneae; 5e et derniére partie. Paris, 6: 979-1298.

Vugdelic, M. 2006 Genetic relations within and amongst Dolomedes aquatic spiders. PhD Thesis, University of East Anglia.

Vugdelic, M., Goodacre, S., Smith H. & Hewitt, G. 2003  Preliminary analysis of the genetic structure in the fen raft spider Dolomedes plantarius (Araneae: Pisauridae). European Arachnology (2003) (Eds. Logunov, D.V. & Penney, D.), 343-348.


Distribution and conservation of other Dolomedes species





  Literature on other Pisaurids

Austad, S.N. & Thornhill, R. 1986  Female reproductive variation in a nuptial-feeding spider Pisaura mirabilis. Bull. Br. Arachnol. 7, 48–52.

Bilde1, T., Tuni1, C., Elsayed1, R., Pekár1, S. & Toft1, S. 2006  Death feigning in the face of sexual cannibalism. Biology Letters, 2, (1), 23-25.

Dondale, C.D, Legendre, R. 1971  Winter diapause in a Mediterranean population of Pisaura mirabilis (Clerck). Bull. Brit. Arachnol. Soc.

Drengsgaard, IL, Toft, S. 1999  Sperm competition in a nuptial feeding spider, Pisaura mirabilis Behaviour, 136, (7), 877-897.

Frost, C., 2005 Ballooning in Dolomedes triton. Canadian Entomolgist 6, 7–9.

Hansen, L. S., Gonzalez,S. F., Toft, S. & Bilde, T 2008  Thanatosis as an adaptive male mating strategy in the nuptial gift-giving spider Pisaura mirabilis Behav. Ecol., 165 (2).

Lenler-Eriksen, P. 1969  The hunting-web of the young spider Pisaura mirabilis J. Zool.

Nentwig, W. 1986  Non-web building spiders: prey specialists or generalists? Oecologia, 69, (4), 571-576

Nentwig, W. & Wissel, C. 1986  A comparison of prey lengths among spiders. Oecologia, 68, (4), 595-600.

Stalhandske, P.  2001  Nuptial gift in the spider Pisaura mirabilis maintained by sexual selection. Behavioral Ecology Vol. 12 No. 6: 691-697


  Key general references for spider identification and general biology

Beccaloni, J. 2009  Arachnids. Natural History Museum, London.

Bristowe, W.S. 1958  The World of Spiders (New Naturalist Series 38) Collins, London

Platnik, N. 2008 The world spider catalog, version 8.0 click here

Roberts, M.J. 1995  The Spiders of Great Britain and Ireland (3 Vols.). Hartley Books, Colchester.

Roberts, M.J. 1995  Field Guide to Spiders of Britain and Northern Europe, HarperCollins, London